Yes, I'm making that 'assumption' because the article says, "... open-access journals, such as PLoS ONE, which charge authors a handling fee to cover their operating costs ..."
EDIT: Link from comment above shows fees for the PLoS family of journals start at $1350 per article and go up rapidly from there.
I absolutely agree. The pay-for-publishing model is heavily disadvantaging towards smaller research institutions, independent researchers or young researchers.
EDIT: Link from comment above shows fees for the PLoS family of journals start at $1350 per article and go up rapidly from there.
http://www.plos.org/journals/pubfees.php