I think that's the "faulty premise" that TheSpiceOfLife is referring to. He's saying that building a study to explain cooperation between humans is modeling a phenomena that's a social construct anyway, when the real question is "what genes lead to behavior that ensures the survival and propagation of those genes?" Natural selection operates on the genetic level, not on the human level, so Kümmerli made an error by trying to inject natural selection arguments into a model where the primary actors are humans.
